The animal was born with GFP so the brain would have adapted to it being present. Quite different from having GFP introduced into an adult retina.
"Less fit" is relative to the context! In a controlled environment like a fish tank I doubt it would be less fit - out in the wild might be different.
I believe this is the paper that describes that specific transgenic axolotl. It is not clear which transgene was used.
http://www.sciencedirect.com/science/article/pii/S0012160605008638
Cheers,
Scott
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